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Perona- One Piece
Username for this character- *GhostPrincess* **Information from Fandom.com**
"Ghost Princess" Perona[7] is the former commander of the Wild Zombies and Surprise Zombies of Thriller Bark prior to its collapse.[3] She was one of the Mysterious Four[2] and one of the major antagonists of the Thriller Bark Saga.
After the defeat of Gecko Moria's army and during the timeskip, she begrudgingly helped[8] and cooperated with Roronoa Zoro while in Dracule Mihawk's home on Kuraigana Island.[9]
Perona has a very young, childlike and immature personality; frequently acting selfishly and on her own whims.[14][15] She dislikes people telling her what to do[16] and disobeying her will.[17] Despite this, she is normally loyal to Gecko Moria[18] and apparently sees him as a father-like role model, as she was crying to rejoin him after being sent to Kuraigana Island.[19] After Zoro appeared on Kuraigana as well, she tended to his wounds, displaying a willingness to befriend even those she formerly knew as a foe and demonstrating that she can be kind to others, despite her normally cruel demeanor.[8]
She is very fond of cute things and will only allow someone to work for her if they are cute.[20] Her dream is to create a kingdom where all the cute things in the world have been turned into zombie animals that obey only her.[21] She dislikes Kumashi because, although he is cute in appearance, his voice is not, and thus she prefers him to just be quiet.[22] Her desire for cute things gives her a slightly childish outlook on some issues.[20] She is somewhat playful during combat, threatening to crush Usopp's heart when she was in her ghost form, which nearly caused Usopp to die from fear, and shortly afterward merely laughing it off when revealing that she could not do so.[23]
Perona has katsaridaphobia, a fear of cockroaches, to the point of screaming and crying when Usopp covered her with fake ones.[24] She is also somewhat naive, having watched Usopp seemingly lift a 10-ton hammer without questioning whether or not his feat was genuine, and shortly afterward passing out from fear when he attacked her with it, despite not actually being harmed.[25]
She is very confident in her Devil Fruit powers being able to defeat anyone.[26] Even when confronted by the powerful and dreaded Warlord of the Sea Bartholomew Kuma, after initially being frozen with fear, Perona quickly regained her confidence and proclaimed that she would defeat him.[27] She is also quite stubborn about using them in combat, having continued to use her Negative Hollows against Usopp even after they proved useless due to his negativity because she believed that his immunity was a fluke.[28]
After the timeskip, she has become more mature, although she still carries a teddy bear around, indicating that she is still obsessed with cuteness to some degree.[12] She displayed a grateful side, as she tearfully thanked Dracule Mihawk for letting her stay on his island.[29]
Like many others, Perona has her own unique laugh, which uses the word Horo (e.g., "Horo-horo-horo-horo-horo!"); this is based on the name of her Devil Fruit, the Horo Horo no Mi.[30]
Relationships:
Thriller Bark Pirates---
Perona followed her master, Gecko Moria, for at least ten years.[11] Her loyalty to him is somewhat complex: when she sensed defeat, she attempted to leave and abandon him[15] while noting that she had only stayed with him for fun,[31] but tried to take Bartholomew Kuma down as a final gift for Moria.[32] Despite not clearly showing it, she is said to love Moria like a father since he found her and raised her as a child, and when the newspaper falsely reported that Moria died in the Whitebeard War, Perona was deeply saddened.[33][34] When she learned that Moria is alive two years later, Perona is overjoyed, and she quickly departed Mihawk's castle to go find him.[35]
Perona had a mixed relationship with her teddy bear servant, Kumashi. While she considered him to be cute in appearance, she forbade him from speaking because his voice is not cute.[22] However, she was devastated when Usopp purified Kumashi.[36]
Strawhats---
While on Thriller Bark, Perona became enemies with one particular member of the Straw Hat Pirates, Usopp, due to the latter becoming the first person to ever resist the effects of her Negative Hollows[37] and later defeating her in a humiliating manner.[25] However, she also cannot help but pity Usopp who lives his life in negativity and attempts to cheer him up, though this may be for her own benefit, as she cannot make him negative unless he was already slightly positive.[38][39] She looked down on Nami when the latter confronted her later, claiming that she could easily defeat Nami with her Mini Hollows because Usopp was her only natural enemy.[40]
After encountering Zoro on Kuraigana Island, she was shocked to see him but decided to treat his wounds as he was the only other person there at the time.[8] Despite their initial quarrels,[41] the two developed a relationship of mutual trust by living on the island for two years alongside Dracule Mihawk.[42] She also willingly brought Zoro back to Sabaody Archipelago[43] and later held back the Marines in order to allow the Straw Hats to escape.[44]
Dracule Mihawk---
After being sent to Kuraigana Island by Kuma's abilities, Perona decided to take residence in the castle where the at the time Warlord Mihawk lived.[45] Mihawk seemingly agreed or simply did not mind letting her stay with him, though he did not display any hospitality toward her beyond that.[46]
It was shown in From the Decks of the World: The 500,000,000 Man Arc that even after going back to Thriller Bark to retrieve Kumashi, she returned to Kuraigana Island and was seen helping him with his work in tilling the land, suggesting a good status between them.[47] When she learned of her old master's survival, she was upset that Mihawk did not tell her and nearly left as he told her to be safe while she tearfully thanked him for letting her stay with him.[35]
Powers/Abilities---
Being formerly one of the Mysterious Four, Perona had a number of zombies under her command prior to her defection from Thriller Bark.[3] Her ex-crewmate, Doctor Hogback, acknowledges her value under Moria's employment.[48]
Perona is physically strong, as she was seen carrying Kumashi's body, which is bigger than she is, while levitating with her Devil Fruit powers.
Perona is able to cook, having cooked several times during the two years while she was living with Mihawk.[35]
- Medical Knowledge:
While she seems to recognize that bandages can help a person who is wounded, she seems to lack expertise in medicine. During the process of treating Zoro's wounds, she covered him from head to toe in an overabundance of bandages the first time she found him.[8] However, despite this fault, she has common sense or enough knowledge of medical skills, as Zoro recovered from his injuries.[49]
-Navigation Knowledge:
She is a capable navigator, demonstrated when she brought Zoro to Sabaody Archipelago all the way from Kuraigana Island.[43] She later traveled to Thriller Bark and returned back to Kuraigana Island all by herself. After learning Moria was alive, she traveled all the way to Hachinosu from Kuraigana Island, going from Paradise to the New World.
-Devil Fruit:
Perona ate the Horo Horo no Mi, a Paramecia-type Devil Fruit that allows her to create and control ghosts. The ghosts have the power to drain the morale, self-esteem, and will to live of anyone whom they pass through, described as hollowing out the victim's spirit.[50][51] She can also create smaller ghosts that explode with enough force to damage stone walls[52] and larger ghosts that produce an even more powerful blast.[53]
Her ghosts can also be used to form a "Ghost Network" for surveillance. This allows her to spy on enemies.[54][55] The ghosts can also duplicate themselves into even more ghosts.[56]
In addition to creating ghosts, she can make her own ghost leave her body. Her disembodied ghost is completely intangible; she can pass through solid objects and cannot be hit by enemy attacks, although she cannot damage anything or attack her opponents physically.[57][58] Her ghost also has the power to levitate and can change the size of her spectral body.[59] However, while she is doing this, her true body remains
Past:
When she was a child, Perona owned a pet bear named Kumae.[10] At some point in her childhood, she met the pirate Gecko Moria who took her in and raised her and she eventually saw him as a father figure.[34]
Over 10 years ago, she joined Moria's crew merely for fun.[31] She and Absalom were present when Moria met and recruited the genius doctor, Hogback.[11]
Five years ago, she was sleeping while Brook was causing trouble in Thriller Bark, which would later cause her to not know what he looked like when he returned.[61] When Kumashi tried to wake her up, she simply told him to shut up and went back to sleep.[62] Three and a half years ago, Perona came to Moria with news that Bartholomew Kuma was elected the new Warlord of the Sea, Moria brushed it off finding his moniker funny but Perona told him that he must be a strong figure to be chosen as a Warlord.[63]
Thriller Bark Saga
Thriller Bark Arc
Perona used her Ghost Network to scout out the Thousand Sunny[1] and later reported the crew's bounties to Hogback and Absalom.[64]
In the Dead Forest, a group of Negative Hollows encountered some of the Straw Hats and, after evading capture, demoralized Monkey D. Luffy, Franky, and Roronoa Zoro.[65] As the Night Attack began, Perona recalled her ghosts and rallied her Wild Zombies and Surprise Zombies.[66] When she left to oversee the looting of the Thousand Sunny, she ordered Kumashi to meet the Risky Brothers, who were carrying Nami, Usopp, and Tony Tony Chopper in coffins, and take them to the Dance Hall.[67]
On the Thousand Sunny, Perona was frustrated at the lack of treasure and had the food taken from the ship. Hildon reported to Perona that Moria was gathering the other members of the Mysterious Four because he had captured Luffy and would be debuting Zombie 900, much to her surprise. Soon afterward, Perona returned to her room, where she found Absalom speaking with Kumashi about the escape of Nami, Usopp, and Chopper. After scolding Kumashi for trying to speak, Perona informed Absalom that Moria was calling them together and explained why he was doing so.[68] In the Mast Mansion, the Mysterious Four discussed Luffy and the Straw Hats, and Perona reported that Nami, Usopp, and Chopper had escaped before yelling at Kumashi when he tried to correct her. When Luffy broke free from his cage, Perona quickly stopped him with her Negative Hollow before watching on as Moria removed Luffy's shadow.[69] After hearing Nami, Usopp, and Chopper talking inside Kumashi and believing that it was Kumashi himself doing so, Perona proclaimed that her dream of creating a kingdom full of cute subordinates was closer to fruition with Luffy's shadow extracted. When two Spider Mice appeared and reported that Tararan's shadow had been extracted, Hogback noted that it could only have been the "Humming Swordsman", a name which Perona did not recognize. Soon afterward, Perona accompanied the rest of the Mysterious Four to the enormous freezer containing the corpse of Oars.[70] Perona watched as Moria used Luffy's shadow to revive Oars's corpse. When Oars awoke, Nami, Usopp, and Chopper screamed and fell from Kumashi's back, prompting Perona to yell at Kumashi for hiding the pirates.[71] Perona continued to chastise Kumashi for hiding the pirates without letting him respond.[72]
After reporting Oars's current activities to Moria, Perona learned that the Straw Hats had learned of the zombies' weakness to salt and were approaching them, with the General Zombies preoccupied with attending Absalom's wedding, but dismissed Hogback's worry over the zombies' weakness being known and retired to her room before attacking the Straw Hats with her ghosts, planning to weaken them enough to be taken by Soldier Zombies. However, Perona's attack was interrupted by Oars crashing down through the mansion.[73] After Franky rebuilt the destroyed bridge, he, Zoro, Sanji, and Usopp charged into Perona's room, where Perona confronted them. After explaining the powers granted by her Devil Fruit, Perona sent a Negative Hollow through each of the Straw Hats, but Usopp was unaffected and purified several of her zombies before declaring that he was always negative, shocking Perona, who tried to cheer him up alongside her zombies before preparing to battle him.[38]
Believing Usopp's immunity to have been a fluke, Perona sent four Negative Hollows through him, but instead became depressed herself alongside her ghosts, to the shock of her zombies.[74] Soon afterward, Perona fled from Usopp while noting that she could defeat the other Straw Hats if it were not for him. After Kumashi distracted Usopp by attacking him, Perona projected her spirit from her body after hiding and tricked Usopp into believing she was flying while proclaiming that she had come up with a plan to defeat him.[39] Declaring that she had simply been caught up in the moment before, Perona attempted to cheer Usopp up once more before appearing behind him in a giant form. After dodging Usopp's Ageha Ryusei, Perona reverted to her normal size before placing her hands inside of Usopp's chest and pretending that she was going to crush his heart, only to reveal that she could not do such a thing. Usopp attacked her with Rokuren Mamushi Boshi, but Perona demonstrated her intangibility by letting the attack go through her and moving through the floor. When Usopp pointed out that she could not touch him either and demanded to know how she could attack him, Perona summoned some Mini Hollows and demonstrated their explosive power on a nearby wall before attacking Usopp with one as well. After tricking Usopp into running into a pillar, Perona covered his body with Mini Hollows and detonated all of them before gleefully watching Kumashi brutalize Usopp. However, Perona was shocked to see Usopp, who had donned his Sogeking mask, purify Kumashi with a salt ball before fleeing once more. Wounding Usopp with a barrage of Mini Hollow blasts once more, Perona decided to hand him over to Moria, but was surprised when Usopp figured out her trick and used Atlas Suisei to expose the room containing her true body.[75]
After Usopp deduced how she left her body by becoming a ghost in order to fight him, Perona revealed that she was a fully conscious Hollow instead of merely being a ghost controlled by her body, prompting Usopp to seemingly attack her body in an attempt to destroy it. After the attack apparently missed, a panicked Perona engulfed his body with a Toku Hollow before returning to her own body. Perona detonated the Toku Hollow but was surprised to see the explosion had disappeared as Usopp revealed he had absorbed it with an Impact Dial. After learning that Usopp had actually shot a Torimochi Boshi at her bed in order to trap her body in one place, Perona witnessed Usopp defeat Hippo Gentleman with a blast from his Impact Dial before covering Perona herself in fake cockroaches with Kurobikari Boshi. Crying and screaming hysterically at the sight of cockroaches, a shocked Perona saw Usopp seemingly lifting a 10-ton hammer and begged him to spare her before passing out in fright as the fake hammer burst apart on impact with her. Usopp celebrated his victory over her as Perona collapsed while foaming at the mouth.[76]
Later, Perona's zombies found Perona still unconscious but mostly unharmed as Oars rampaged through Thriller Bark.[77] Soon afterward, Perona regained consciousness and had to be calmed down by her zombies when she acted like Usopp was still attacking her. Upon seeing the damage done to the mansion by Oars, Perona wondered if the Straw Hats had done this and was shocked to learn that Oars had completely wiped out the General Zombies as well, which caused her to question if they could control him at all. However, the zombies' explanation of what was happening was interrupted by Oars punching them through one of the nearby walls while searching for Nami; witnessing this, Perona was convinced that Oars and the Straw Hats were going to completely destroy Thriller Bark and ordered her remaining subordinates to gather all the treasure and some supplies so they could leave on the Thousand Sunny.[78]
Observing the battle between the Straw Hats and Oars from the Thousand Sunny, Perona admitted that she only joined Moria for fun and would not stay with him if her life were in danger before telling her subordinates that they would be leaving without delay.[79] When Nami confronted her over her taking the treasure and the Thousand Sunny, Perona declared that she was planning to leave Thriller Bark on the Thousand Sunny and prepared to battle while proclaiming that her Mini Hollows should be enough to defeat Nami. However, she was interrupted by the appearance of Bartholomew Kuma, whom Perona was terrified at the sight of due to knowing his reputation as a cruel pirate and a Warlord of the Sea. When Kuma asked her where she would like to go on a trip, Perona described a dark, damp island with an old castle filled with malice before realizing that she had gotten caught up in his pace. Resolving to take down Kuma as a parting gift for Moria due to his shadow likely being strong, Perona prepared to attack him with a combination of Negative Hollows and a Toku Hollow, but Kuma suddenly made her disappear before her attack could connect, to the confusion of those watching.[80]
When Hogback and Absalom escaped Thriller Bark later, they left without Perona while noting they could not find her anywhere.[81]
***To see more about her past, visit this site: onepiece.fandom.com/wiki/Perona#Personality***
"Ghost Princess" Perona[7] is the former commander of the Wild Zombies and Surprise Zombies of Thriller Bark prior to its collapse.[3] She was one of the Mysterious Four[2] and one of the major antagonists of the Thriller Bark Saga.
After the defeat of Gecko Moria's army and during the timeskip, she begrudgingly helped[8] and cooperated with Roronoa Zoro while in Dracule Mihawk's home on Kuraigana Island.[9]
Perona has a very young, childlike and immature personality; frequently acting selfishly and on her own whims.[14][15] She dislikes people telling her what to do[16] and disobeying her will.[17] Despite this, she is normally loyal to Gecko Moria[18] and apparently sees him as a father-like role model, as she was crying to rejoin him after being sent to Kuraigana Island.[19] After Zoro appeared on Kuraigana as well, she tended to his wounds, displaying a willingness to befriend even those she formerly knew as a foe and demonstrating that she can be kind to others, despite her normally cruel demeanor.[8]
She is very fond of cute things and will only allow someone to work for her if they are cute.[20] Her dream is to create a kingdom where all the cute things in the world have been turned into zombie animals that obey only her.[21] She dislikes Kumashi because, although he is cute in appearance, his voice is not, and thus she prefers him to just be quiet.[22] Her desire for cute things gives her a slightly childish outlook on some issues.[20] She is somewhat playful during combat, threatening to crush Usopp's heart when she was in her ghost form, which nearly caused Usopp to die from fear, and shortly afterward merely laughing it off when revealing that she could not do so.[23]
Perona has katsaridaphobia, a fear of cockroaches, to the point of screaming and crying when Usopp covered her with fake ones.[24] She is also somewhat naive, having watched Usopp seemingly lift a 10-ton hammer without questioning whether or not his feat was genuine, and shortly afterward passing out from fear when he attacked her with it, despite not actually being harmed.[25]
She is very confident in her Devil Fruit powers being able to defeat anyone.[26] Even when confronted by the powerful and dreaded Warlord of the Sea Bartholomew Kuma, after initially being frozen with fear, Perona quickly regained her confidence and proclaimed that she would defeat him.[27] She is also quite stubborn about using them in combat, having continued to use her Negative Hollows against Usopp even after they proved useless due to his negativity because she believed that his immunity was a fluke.[28]
After the timeskip, she has become more mature, although she still carries a teddy bear around, indicating that she is still obsessed with cuteness to some degree.[12] She displayed a grateful side, as she tearfully thanked Dracule Mihawk for letting her stay on his island.[29]
Like many others, Perona has her own unique laugh, which uses the word Horo (e.g., "Horo-horo-horo-horo-horo!"); this is based on the name of her Devil Fruit, the Horo Horo no Mi.[30]
Relationships:
Thriller Bark Pirates---
Perona followed her master, Gecko Moria, for at least ten years.[11] Her loyalty to him is somewhat complex: when she sensed defeat, she attempted to leave and abandon him[15] while noting that she had only stayed with him for fun,[31] but tried to take Bartholomew Kuma down as a final gift for Moria.[32] Despite not clearly showing it, she is said to love Moria like a father since he found her and raised her as a child, and when the newspaper falsely reported that Moria died in the Whitebeard War, Perona was deeply saddened.[33][34] When she learned that Moria is alive two years later, Perona is overjoyed, and she quickly departed Mihawk's castle to go find him.[35]
Perona had a mixed relationship with her teddy bear servant, Kumashi. While she considered him to be cute in appearance, she forbade him from speaking because his voice is not cute.[22] However, she was devastated when Usopp purified Kumashi.[36]
Strawhats---
While on Thriller Bark, Perona became enemies with one particular member of the Straw Hat Pirates, Usopp, due to the latter becoming the first person to ever resist the effects of her Negative Hollows[37] and later defeating her in a humiliating manner.[25] However, she also cannot help but pity Usopp who lives his life in negativity and attempts to cheer him up, though this may be for her own benefit, as she cannot make him negative unless he was already slightly positive.[38][39] She looked down on Nami when the latter confronted her later, claiming that she could easily defeat Nami with her Mini Hollows because Usopp was her only natural enemy.[40]
After encountering Zoro on Kuraigana Island, she was shocked to see him but decided to treat his wounds as he was the only other person there at the time.[8] Despite their initial quarrels,[41] the two developed a relationship of mutual trust by living on the island for two years alongside Dracule Mihawk.[42] She also willingly brought Zoro back to Sabaody Archipelago[43] and later held back the Marines in order to allow the Straw Hats to escape.[44]
Dracule Mihawk---
After being sent to Kuraigana Island by Kuma's abilities, Perona decided to take residence in the castle where the at the time Warlord Mihawk lived.[45] Mihawk seemingly agreed or simply did not mind letting her stay with him, though he did not display any hospitality toward her beyond that.[46]
It was shown in From the Decks of the World: The 500,000,000 Man Arc that even after going back to Thriller Bark to retrieve Kumashi, she returned to Kuraigana Island and was seen helping him with his work in tilling the land, suggesting a good status between them.[47] When she learned of her old master's survival, she was upset that Mihawk did not tell her and nearly left as he told her to be safe while she tearfully thanked him for letting her stay with him.[35]
Powers/Abilities---
Being formerly one of the Mysterious Four, Perona had a number of zombies under her command prior to her defection from Thriller Bark.[3] Her ex-crewmate, Doctor Hogback, acknowledges her value under Moria's employment.[48]
Perona is physically strong, as she was seen carrying Kumashi's body, which is bigger than she is, while levitating with her Devil Fruit powers.
Perona is able to cook, having cooked several times during the two years while she was living with Mihawk.[35]
- Medical Knowledge:
While she seems to recognize that bandages can help a person who is wounded, she seems to lack expertise in medicine. During the process of treating Zoro's wounds, she covered him from head to toe in an overabundance of bandages the first time she found him.[8] However, despite this fault, she has common sense or enough knowledge of medical skills, as Zoro recovered from his injuries.[49]
-Navigation Knowledge:
She is a capable navigator, demonstrated when she brought Zoro to Sabaody Archipelago all the way from Kuraigana Island.[43] She later traveled to Thriller Bark and returned back to Kuraigana Island all by herself. After learning Moria was alive, she traveled all the way to Hachinosu from Kuraigana Island, going from Paradise to the New World.
-Devil Fruit:
Perona ate the Horo Horo no Mi, a Paramecia-type Devil Fruit that allows her to create and control ghosts. The ghosts have the power to drain the morale, self-esteem, and will to live of anyone whom they pass through, described as hollowing out the victim's spirit.[50][51] She can also create smaller ghosts that explode with enough force to damage stone walls[52] and larger ghosts that produce an even more powerful blast.[53]
Her ghosts can also be used to form a "Ghost Network" for surveillance. This allows her to spy on enemies.[54][55] The ghosts can also duplicate themselves into even more ghosts.[56]
In addition to creating ghosts, she can make her own ghost leave her body. Her disembodied ghost is completely intangible; she can pass through solid objects and cannot be hit by enemy attacks, although she cannot damage anything or attack her opponents physically.[57][58] Her ghost also has the power to levitate and can change the size of her spectral body.[59] However, while she is doing this, her true body remains
Past:
When she was a child, Perona owned a pet bear named Kumae.[10] At some point in her childhood, she met the pirate Gecko Moria who took her in and raised her and she eventually saw him as a father figure.[34]
Over 10 years ago, she joined Moria's crew merely for fun.[31] She and Absalom were present when Moria met and recruited the genius doctor, Hogback.[11]
Five years ago, she was sleeping while Brook was causing trouble in Thriller Bark, which would later cause her to not know what he looked like when he returned.[61] When Kumashi tried to wake her up, she simply told him to shut up and went back to sleep.[62] Three and a half years ago, Perona came to Moria with news that Bartholomew Kuma was elected the new Warlord of the Sea, Moria brushed it off finding his moniker funny but Perona told him that he must be a strong figure to be chosen as a Warlord.[63]
Thriller Bark Saga
Thriller Bark Arc
Perona used her Ghost Network to scout out the Thousand Sunny[1] and later reported the crew's bounties to Hogback and Absalom.[64]
In the Dead Forest, a group of Negative Hollows encountered some of the Straw Hats and, after evading capture, demoralized Monkey D. Luffy, Franky, and Roronoa Zoro.[65] As the Night Attack began, Perona recalled her ghosts and rallied her Wild Zombies and Surprise Zombies.[66] When she left to oversee the looting of the Thousand Sunny, she ordered Kumashi to meet the Risky Brothers, who were carrying Nami, Usopp, and Tony Tony Chopper in coffins, and take them to the Dance Hall.[67]
On the Thousand Sunny, Perona was frustrated at the lack of treasure and had the food taken from the ship. Hildon reported to Perona that Moria was gathering the other members of the Mysterious Four because he had captured Luffy and would be debuting Zombie 900, much to her surprise. Soon afterward, Perona returned to her room, where she found Absalom speaking with Kumashi about the escape of Nami, Usopp, and Chopper. After scolding Kumashi for trying to speak, Perona informed Absalom that Moria was calling them together and explained why he was doing so.[68] In the Mast Mansion, the Mysterious Four discussed Luffy and the Straw Hats, and Perona reported that Nami, Usopp, and Chopper had escaped before yelling at Kumashi when he tried to correct her. When Luffy broke free from his cage, Perona quickly stopped him with her Negative Hollow before watching on as Moria removed Luffy's shadow.[69] After hearing Nami, Usopp, and Chopper talking inside Kumashi and believing that it was Kumashi himself doing so, Perona proclaimed that her dream of creating a kingdom full of cute subordinates was closer to fruition with Luffy's shadow extracted. When two Spider Mice appeared and reported that Tararan's shadow had been extracted, Hogback noted that it could only have been the "Humming Swordsman", a name which Perona did not recognize. Soon afterward, Perona accompanied the rest of the Mysterious Four to the enormous freezer containing the corpse of Oars.[70] Perona watched as Moria used Luffy's shadow to revive Oars's corpse. When Oars awoke, Nami, Usopp, and Chopper screamed and fell from Kumashi's back, prompting Perona to yell at Kumashi for hiding the pirates.[71] Perona continued to chastise Kumashi for hiding the pirates without letting him respond.[72]
After reporting Oars's current activities to Moria, Perona learned that the Straw Hats had learned of the zombies' weakness to salt and were approaching them, with the General Zombies preoccupied with attending Absalom's wedding, but dismissed Hogback's worry over the zombies' weakness being known and retired to her room before attacking the Straw Hats with her ghosts, planning to weaken them enough to be taken by Soldier Zombies. However, Perona's attack was interrupted by Oars crashing down through the mansion.[73] After Franky rebuilt the destroyed bridge, he, Zoro, Sanji, and Usopp charged into Perona's room, where Perona confronted them. After explaining the powers granted by her Devil Fruit, Perona sent a Negative Hollow through each of the Straw Hats, but Usopp was unaffected and purified several of her zombies before declaring that he was always negative, shocking Perona, who tried to cheer him up alongside her zombies before preparing to battle him.[38]
Believing Usopp's immunity to have been a fluke, Perona sent four Negative Hollows through him, but instead became depressed herself alongside her ghosts, to the shock of her zombies.[74] Soon afterward, Perona fled from Usopp while noting that she could defeat the other Straw Hats if it were not for him. After Kumashi distracted Usopp by attacking him, Perona projected her spirit from her body after hiding and tricked Usopp into believing she was flying while proclaiming that she had come up with a plan to defeat him.[39] Declaring that she had simply been caught up in the moment before, Perona attempted to cheer Usopp up once more before appearing behind him in a giant form. After dodging Usopp's Ageha Ryusei, Perona reverted to her normal size before placing her hands inside of Usopp's chest and pretending that she was going to crush his heart, only to reveal that she could not do such a thing. Usopp attacked her with Rokuren Mamushi Boshi, but Perona demonstrated her intangibility by letting the attack go through her and moving through the floor. When Usopp pointed out that she could not touch him either and demanded to know how she could attack him, Perona summoned some Mini Hollows and demonstrated their explosive power on a nearby wall before attacking Usopp with one as well. After tricking Usopp into running into a pillar, Perona covered his body with Mini Hollows and detonated all of them before gleefully watching Kumashi brutalize Usopp. However, Perona was shocked to see Usopp, who had donned his Sogeking mask, purify Kumashi with a salt ball before fleeing once more. Wounding Usopp with a barrage of Mini Hollow blasts once more, Perona decided to hand him over to Moria, but was surprised when Usopp figured out her trick and used Atlas Suisei to expose the room containing her true body.[75]
After Usopp deduced how she left her body by becoming a ghost in order to fight him, Perona revealed that she was a fully conscious Hollow instead of merely being a ghost controlled by her body, prompting Usopp to seemingly attack her body in an attempt to destroy it. After the attack apparently missed, a panicked Perona engulfed his body with a Toku Hollow before returning to her own body. Perona detonated the Toku Hollow but was surprised to see the explosion had disappeared as Usopp revealed he had absorbed it with an Impact Dial. After learning that Usopp had actually shot a Torimochi Boshi at her bed in order to trap her body in one place, Perona witnessed Usopp defeat Hippo Gentleman with a blast from his Impact Dial before covering Perona herself in fake cockroaches with Kurobikari Boshi. Crying and screaming hysterically at the sight of cockroaches, a shocked Perona saw Usopp seemingly lifting a 10-ton hammer and begged him to spare her before passing out in fright as the fake hammer burst apart on impact with her. Usopp celebrated his victory over her as Perona collapsed while foaming at the mouth.[76]
Later, Perona's zombies found Perona still unconscious but mostly unharmed as Oars rampaged through Thriller Bark.[77] Soon afterward, Perona regained consciousness and had to be calmed down by her zombies when she acted like Usopp was still attacking her. Upon seeing the damage done to the mansion by Oars, Perona wondered if the Straw Hats had done this and was shocked to learn that Oars had completely wiped out the General Zombies as well, which caused her to question if they could control him at all. However, the zombies' explanation of what was happening was interrupted by Oars punching them through one of the nearby walls while searching for Nami; witnessing this, Perona was convinced that Oars and the Straw Hats were going to completely destroy Thriller Bark and ordered her remaining subordinates to gather all the treasure and some supplies so they could leave on the Thousand Sunny.[78]
Observing the battle between the Straw Hats and Oars from the Thousand Sunny, Perona admitted that she only joined Moria for fun and would not stay with him if her life were in danger before telling her subordinates that they would be leaving without delay.[79] When Nami confronted her over her taking the treasure and the Thousand Sunny, Perona declared that she was planning to leave Thriller Bark on the Thousand Sunny and prepared to battle while proclaiming that her Mini Hollows should be enough to defeat Nami. However, she was interrupted by the appearance of Bartholomew Kuma, whom Perona was terrified at the sight of due to knowing his reputation as a cruel pirate and a Warlord of the Sea. When Kuma asked her where she would like to go on a trip, Perona described a dark, damp island with an old castle filled with malice before realizing that she had gotten caught up in his pace. Resolving to take down Kuma as a parting gift for Moria due to his shadow likely being strong, Perona prepared to attack him with a combination of Negative Hollows and a Toku Hollow, but Kuma suddenly made her disappear before her attack could connect, to the confusion of those watching.[80]
When Hogback and Absalom escaped Thriller Bark later, they left without Perona while noting they could not find her anywhere.[81]
***To see more about her past, visit this site: onepiece.fandom.com/wiki/Perona#Personality***
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